Thorns deal a percentage of an enemy's max health as damage when they hit your tower (50% damage to bosses). This also includes ranged attackers.
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If you have a Wall, you will also need to take into your Wall Thorns to calculate the amount of enemy hits taken until an enemy dies. You may also refer to Infographic: Wall Thorns Max Hits with Plasma Cannon Mastery & Thorn Relic.
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Consists of 99 levels with each level increasing Thorns with 1% for a total of 99% Thorns.